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Selected readings

There are many published research articles on the amphibians of Borneo and potentially useful literature with broader scope. In this section we recommend some. This is a very subjective list and you may want to expand that list by your own literature reseach. This section is not meant to give a complete compilation of the literature on the frogs of Borneo.


Inger, R.F., Stuebing, R.B., Grafe, T.U. & Dehling, J.M. (2017) A field guide to the frogs of Borneo. 3rd Edition. Natural History Books.

Haas, A., Das, I., Hertwig, S., Schulz-Schaeffer, R. & Bublies, P. (2022) A Guide to the Tadpoles of Borneo. Tredition, Hamburg, 279 pp.


Zug G. R.  (2015) Morphology and Systematics of Kalophynus interlineatus-pleurostigma populations (Anura: Microhylidae: Kalphryninae) and a taxonomy of the genus Kalophrynus Tschudi, Asian Sticky Frogs. Proceedings of the California Academy of Sciences 62:135–190.

Dehling J.M. (2015) A new species of Rhacophorus (Anura: Rhacophoridae) from Gunung Kinabalu, Borneo. Salamandra 51:1–11.

Oberhummer, E., Barten, C., Schweizer, M., Das, I., Haas, A., Hertwig, S.T., 2015. Description of the tadpoles of three rare species of megophryid frogs (Amphibia: Anura: Megophryidae) from Gunung Mulu, Sarawak, Malaysia. Zootaxa, 3835 (1): 059 – 079.

Haas, A., J. Pohlmeyer, D. S. McLeod, T. Kleinteich, S. T. Hertwig, I. Das, D. R. Buchholz, 2014. Extreme tadpoles II: the highly derived larval anatomy of Occidozyga baluensis (Boulenger, 1896), an obligate carnivorous tadpole. Zoomorphology, 133 (3), 321-342

Hertwig, S.T.,  Min, P.Y., Haas, A., Das, I. (2014) Dressed in black: A New Ansonia Stoliczka, 1870 (Lissamphibia: Anura: Bufonidae) from Gunung Murud, Sarawak, East Malaysia (Borneo).
Zootaxa 3814:419–431

Das, I., Min, P.Y., Hsu, W.W., Hertwig, S.T., Haas, A (2014) Red Hot Chilli Pepper. A New Calluella Stoliczka, 1872 (Lissamphibia: Anura: Microhylidae) from Sarawak, East Malaysia (Borneo)
Zootaxa 3785:550–560

Matsui M., Nishikawa K., Eto K. (2014) A new burrow-utilising fanged frog from Sarawak, East Malaysia (Anura: Dicroglossidae). Raffles Bulletin of Zoology 62:679–687.

Matsui M., Zainudin R., Nishikawa K. (2014) A New Species of Leptolalax from Sarawak, Western Borneo (Anura: Megophryidae). ZOO SCI 31:773–779. doi: 10.2108/zs140137

Matsui M., Nishikawa K. (2014) Description of a new species of Limnonectes from Sarawak Malaysian Borneo (Dicroglossidae, Anura). Current Herpetology 33:135–147. doi: 10.5358/hsj.33.135

Dehling J.M., Dehling D.M. (2013) A new montane species of Philautus (Amphibia: Anura: Rhacophroidae) from western Sarawak, Malaysia, Borneo. Zootaxa 3686:277. doi: 10.11646/zootaxa.3686.2.8

Pui, Y.M., Das, I., Haas, A. (2013) New records of Limnonectes rhacodus (Inger, Boeadi and Taufik, 1996) (Lissamphibia: Anura: Dicroglossidae) from Sarawak, East Malaysia (Northwestern Borneo)
Check List 9(6):1588–1589

Hertwig, S. T., Schweizer, M., Das, I., Haas, A. (2013) Diversification in a biodiversity hotspot – the evolution of Southeast Asian rhacophorid tree frogs on Borneo (Amphibia: Anura: Rhacophoridae).
 Molecular Phylogenetics and Evolution 68(3):567–581

Matsui M., Nishikawa K., Yambun P. (2013a) A new Leptolalax from the mountains of Sabah, Borneo (Amphibia, Anura, Megophryidae). Zootaxa 3753:440–452. doi: 10.11646/zootaxa.3753.5.3

Matsui M, Shimada T, Sudin A (2013) A new gliding frog of the genus Rhacophorus from Borneo. Current Herpetol 32:112–124.

Hertwig, S., Lilje, K.E., Min, P.Y., Haas, A., Das, I. (2012) Molecular evidence of direct development in the rhacophorid frog Philautus acutus (Rhacophroidae, anura) from Borneo.
Raffles Bulletin of Zoology 60:555–563

Haas, A., Hertwig, S. T., Krings, W., Braskamp, E., Dehling, J. M., Min, P. Y., Jankowski, A., Schweizer, M. Das, I. (2012) Description of three Rhacophorus tadpoles (Lissamphibia: Anura: Rhacophoridae) from Sarawak, Malaysia (Borneo).
Zootaxa 3328:1–19

Hamidy A, Matsui M, Nishikawa K, Belabut DM (2012) Detection of cryptic taxa in Leptobrachium nigrops (Amphibia, Anura, Megophryidae), with description of two new species. Zootaxa 22–39.

Hertwig, S.T., Das, I., Schweizer, M., Brown, R., Has, A. (2012): Phylogenetic relationships of the Rhacophorus everetti-group and implications for the evolution of reproductive modes in Philautus (Amphibia: Anura: Rhacophoridae). Zoologica Scripta, 41 (1), 29–46.

Wildenhues, M., Rauhaus, A., Bach, R., Karbe, D., van der Straeten, K., Hertwig, S.T., Ziegler, T. (2012): Husbandry, captive breeding, larval development and stages of the Malayan horned frog Megophrys nasuta (Schlegel, 1858) (Amphibia: Anura: Megophryidae). Amphibian and Reptile Conservation 5(3):15-28.

Das, I. & Haas, A. (2010) New species of Microhyla from Sarawak: Old World’s smallest frogs crawl out of miniature pitcher plants on Borneo (Amphibia: Anura: Microhylidae). Zootaxa, 2571: 37–52.

Haas, A., Wolter, J., Hertwig, S.T., Das, I. (2009): Larval morphologies of three species of stream toads, genus Ansonia (Amphibia: Bufonidae) from East Malaysia (Borneo), with a key to known Bornean Ansonia tadpoles. Zootaxa 2302: 1–18.

Haas, A., Das, A. (2008) Larval identities of Ansonia hanitschi Inger, 1960 (Amphibia: Bufonidae) and Polypedates colletti (Boulenger, 1890) (Amphibia: Rhacophoridae) from East Malaysia (Borneo). Salamandra 44(2):85–100.

Das, I., Jankowski, A., Mohd. Iqbal b. Makmor , Haas, A. 2007. Species diversity, elevational distribution and reproductive modes in an amphibian community at the Matang Range, Sarawak (Borneo). Mitteilungen aus dem Zoologischen Museums Hamburg 104: 141–174.

Handrigan, G. R., A. Haas and R. J. Wassersug. 2007. Bony-tailed tadpoles: the development of supernumerary caudal vertebrae in larval megophryids (Anura). Evol Dev 9: 190-202.

Das, I. and A. Haas. 2006. Sources of larval identities for amphibians from Borneo. Herpetological Review 36: 375-382.

Haas, A., S. Hertwig and I. Das. 2006. Extreme tadpoles: the morphology of the fossorial megophryid larva, Leptobrachella mjobergi. Zoology (Jena) 109: 26-42.

Das, I. and A. Haas. 2005. A new species of Rhacophorus (Anura: Rhacophoridae) from Gunung Gading, Sarawak (Western Borneo). Raffles Bulletin of Zoology 53: 257–263.

Das, I. and A. Haas. 2003. A new species of Kalophrynus (Anura: Microhylidae) from Central Borneo. Raffles Bulletin of Zoology 51: 113–117.

Anstis, M. 2002. The tadpoles of South-Eastern Australia. New Holland Publishers (Australia), Sydney, Auckland, London, Cape Town.

Arifin, U., Iskandar, D., Bickford, D., Meier, R., Kutty, S. N., et al. (2011). Phylogenetic relationships within the genus Staurois (Anura, Ranidae) based on 16S rRNA sequences. Zootaxa, 2744, 39–52.

Chou, W.-h. and J.-Y. Lin. 1997. Tadpoles of Taiwan. Special Publication National Museum of Natural Science 7: 1–98.

Dring, J. 1987 Bornean treefrogs of the genus Philautus (Rhacophoridae). Amphibia-Reptilia 8:19–47.

Inger, R. F. 2009. Contributions to the natural history of seven species of Bornean frogs. Fieldiana Zoology New Seruies 116:1–25.

Inger, R. F., Stuart, B. L., Iskandar, D. T. 2009. Systematics of a widespread Southeast Asian frog, Rana chalconota (Amphibia: Anura: Ranidae). Zoological Journal of the Linnean Society 155: 123–147.

Dehling, M. 2008. A new treefrog (Anura: Rhacophoridae: Rhacophorus) from Gunung Mulu, Borneo. Salamandra 44(4): 193–205.

Dehling, M., Grafe, U. 2008. A new treefrog of the genus Rhacophorus (Anura: Rhacophoridae) from Brunei Darussalam (Borneo). Salamandra 44: 101–112.

Inger, R. F. 1985. Tadpoles of the forested regions of Borneo. Fieldiana Zool. 26: 1-89.

Inger, R. F. and R. B. Stuebing. 2005. A field guide to the frogs of Borneo. Natural History Publications (Borneo), Kota Kinabalu.

Malkmus, R., U. Manthey, G. Vogel, P. Hoffmann and J. Kosuch. 2002. Amphibians and reptiles of Mount Kinabalu (North Borneo). Koeltz Scientific Books, Koenigstein.

Manthey, U. and W. Grossmann. 1997. Amphibien & Reptilien Südostasiens. Natur und Tier-Verlag, Münster.

Matsui, M., Mohamed, M., Shimada, T., & Sudin, A. (2007). Resurrection of Staurois parvus from S. tuberilinguis from Borneo (Amphibia, Ranidae). Zoological Science, 24(1), 101–106.

Matsui, M., Yambun, P., Sudin, A. 2007. Taxonomic relationships of Ansonia anotis Inger, Tan, 

and Yambun, 2001 and Pedostibes maculatus (Mocquard, 1890), with a description of a new genus (Amphibia, Bufonidae). Zoological Science 24: 1159–1166.

Matsui, M., Shimada, T. & Sudin, A. (2010) A new species of Meristogenys (Amphibia, Anura, Ranidae) from Sabah, Borneo. Zoological Science, 27, 61–66.

McDiarmid, R. W. and R. Altig. 1999. Tadpoles. University Chicago Press, Chicago, London.