We thank the following institutions for supporting the project:


Volkswagen Foundation Germany

Swiss Academy of Sciences

Universiti Malaysia Sarawak (UNIMAS)

IBEC (Institute for Biodiversity and Environmental Conservation)

University Malaysia Sabah: ITBC

Universität Hamburg

Natural History Museum Berne

Economic Planning Unit

Sabah Parks

Forest Department of Sarawak

National Parks Staff




The support by our commercial sponsor is gratefully acknowledged:

Zweibrüder Optoelectronics
We gratefully acknowledge the granting of permission to cross-link with:

Amphibia Species of the World by Dr. Darrel Frost, American Museum of Natural History for taxonomic information about species.

and Amphibia Web Species Maps by:

AmphibiaWeb: Information on amphibian biology and conservation. [web application]. 2009. Berkeley, California: AmphibiaWeb. Available: http://amphibiaweb.org/
We wish to thank all colleagues, students, friends, and other individuals who have provided information, images, or have helped us in the field and in the lab (in random order):

Jörg Hofmann, Andreas Sentker, Johanna Wolter, Torben Riehl, Daniela Haarmeyer, Melitta Wunderskirchner, Cindy Hefenbrock, Sebastian Marwitz, Despina Chaluppa, Stine Griep, Helena Dobbeck, Stephan Henne, Felix Meyer, Julia Juchheim, Pui Min Yong, Jeet Sukumaran, Kueh Boon Hee, Sandra James Tinggom, Castro Micheal
Lily Sir, Mohd. Iqbal Makmor, Khairul Anuar bin Omar, Yolande Direp ak Michael Direp, Hairi bin Hedeir, Siti Shuhada bt Mustaffa, Mona Octavia Sulai, Tzi Ming Leong, Frederick Francis Gnathau, Michael Ready, Nick Brown